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Mating may occur outside the rookeries, between non-territorial males and females, as the latter move to and from the mating site.
en.wikipedia.org Breeding takes place in these abandoned nests; usually breeding occurs colonially in rookeries because these birds tend to stay together in groups.
en.wikipedia.org While their rookeries had been thickly buried as well, coastal erosion had cleared them by the summer of 2009.
en.wikipedia.org The mixed riparian forest habitat on the refuge is important for breeding and migrating passerine birds, and supports a large heron/egret rookery.
en.wikipedia.org Thus, it is better not to visit the rookery after a heavy rain fall or during winter and spring rainy seasons.
en.wikipedia.org