He studied marine biology, zoology, botany, and philosophy, and became a well-known entomologist.
en.wikipedia.org He is the author of about 120 papers on international journals of zoology, ethology, marine biology and marine ecology.
en.wikipedia.org He taught courses in evolutionary theory and human origins, zoology and anthropology.
en.wikipedia.org He contributed to the life sciences, especially botany and zoology, and to linguistics, folklore, ethnography, history, literary studies, and archaeology.
en.wikipedia.org For part of this time (1901-1903) he was a conservator in the museum of zoology of the university.
en.wikipedia.org