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ни́жний слой
substratum <-ta> [ˌsʌbˈstrɑ:təm, pl: -tə, америк. ˈsʌbˌstreɪt̬əm, pl: -t̬ə] СУЩ.
substratum
PONS OpenDict

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Одноязычные примеры (не проверены редакцией PONS)
But invasion is the substratum of all such theories even if words like migration are used.
en.wikipedia.org
During metamorphosis, the veliger, depending on species, may secrete an attachment structure called a byssus that anchors it to the substratum.
en.wikipedia.org
Though these "targumim" were later easternized, the substratum belying their origins still remains.
en.wikipedia.org
These are taken as evidence of substratum influence.
en.wikipedia.org
Juvenile specimens or those in the initial phase have a mottled pattern similar to the substratum.
en.wikipedia.org