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подлежащее
Enzym-gekoppelt
enzyme-linked [ˌenzaimˈlɪŋkt] ПРИЛ.
PONS OpenDict

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Одноязычные примеры (не проверены редакцией PONS)
The enzyme-linked secondary antibody follows the detecting antibodies and binds to them.
en.wikipedia.org
The method of detection is through enzyme-linked immunosorbent assay.
en.wikipedia.org
The enzyme acts as an amplifier; even if only few enzyme-linked antibodies remain bound, the enzyme molecules will produce many signal molecules.
en.wikipedia.org
Commercially available activity assays are based on chromogenic assays that use activation by snake venom in an activating reagent, or clotting and enzyme-linked immunosorbant assays.
en.wikipedia.org
However, use of a secondary-antibody conjugate avoids the expensive process of creating enzyme-linked antibodies for every antigen one might want to detect.
en.wikipedia.org